Browsing by Author "Kay, RF"

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    A baseline paleoecological study for the Santa Cruz Formation (late–early Miocene) at the Atlantic coast of Patagonia, Argentina
    (Palaeogeography, Palaeoclimatology, Palaeoecology, 2010-06) Vizcaíno, SF; Bargo, MS; Kay, RF; Fariña, RA; Di Giacomo, M; Perry, JMG; Prevosti, FJ; Toledo, N; Cassini, GH; Fernicola, JC
    Coastal exposures of the Santa Cruz Formation (late-early Miocene, southern Patagonia, Argentina) between the Coyle and Gallegos rivers have been a fertile ground for recovery of Miocene vertebrates for more than 100 years. The formation contains an exceptionally rich mammal fauna, which documents a vertebrate assemblage very different from any living community, even at the ordinal level. Intensive fieldwork performed since 2003 (nearly 1200 specimens have been collected, including marsupials, xenarthrans, notoungulates, litopterns astrapotheres, rodents, and primates) document this assertion. The goal of this study is to attempt to reconstruct the trophic structure of the Santacrucian mammalian community with precise stratigraphic control. Particularly, we evaluate the depauperate carnivoran paleoguild and identify new working hypotheses about this community. A database has been built from about 390 specimens from two localities: Campo Barranca (CB) and Puesto Estancia La Costa (PLC). All species have been classified as herbivore or carnivore, their body masses estimated, and the following parameters estimated: population density, on-crop biomass, metabolic rates, and the primary and secondary productivity. According to our results, this model predicts an imbalance in both CB and PLC faunas which can be seen by comparing the secondary productivity of the ecosystem and the energetic requirements of the carnivores in it. While in CB, the difference between carnivores and herbivores is six-fold, in PLC this difference is smaller, the secondary productivity is still around three times that of the carnivore to herbivore ratio seen today. If both localities are combined, the difference rises to around four-fold in favour of secondary productivity. Finally, several working hypotheses about the Santacrucian mammalian community and the main lineages of herbivores and carnivores are offered. © 2010 Elsevier B.V. All rights reserved.
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    A New Humerus of Homunculus patagonicus, a Stem Platyrrhine from the Santa Cruz Formation (Late Early Miocene), Santa Cruz Province, Argentina
    (Ameghiniana, 2022-01-01) Fleagle, JG; Gladman, JT; Kay, RF
    We describe a well-preserved humerus of Homunculus patagonicus, a stem platyrrhine from the late early Miocene of the Santa Cruz Formation, Santa Cruz Province, Argentina. The distal part of a humerus was collected by Carlos Ameghino and figured in the 19thCentury, but is now lost. Other described postcranial elements, also collected by him include a femur and a partial radius. Comparative observations are made with living and extinct platyrrhines, Oligocene African anthropoids, and extant strepsirrhines. Homunculus patagonicus was a robustly built arboreal quadruped that weighed between 2.2 and 2.6 kg. There is no evidence that the elbow could be fully extended as in living suspensory platyrrhines like Ateles. The medial orientation of the epicondyle suggests that the finger and wrist flexors were not aligned with the long axis of the limb, a distinction from more cursorial monkeys (extant cercopithecoids and the Cuban Pleistocene fossil platyrrhine Paralouatta have retroflexed medial epicondyles). Overall, the morphology is typically platyrrhine although the bone is quite robust. The robustness of the humerus is most comparable to that of early anthropoids from Africa rather than any extant platyrrhine.
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    Adaptive wear-based changes in dental topography associated with atelid (Mammalia: Primates) diets
    (Biological Journal of the Linnean Society, 2018-06-28) Pampush, JD; Spradley, JP; Morse, PE; Griffith, D; Gladman, JT; Gonzales, LA; Kay, RF
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    Additional Vertebrate Remains from the Early Miocene of Kutch, Gujarat
    (Special Publication of the Paleontological Society of India, 2014) Patnaik, R; Milankumar Sharma, K; Mohan, L; Williams, BA; Kay, RF; Chatrath, P
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    An exceptionally well-preserved skeleton of Palaeothentes from the Early Miocene of Patagonia, Argentina: new insights into the anatomy of extinct paucituberculatan marsupials
    (Swiss Journal of Palaeontology, 2014-07) Forasiepi, AM; Sánchez-Villagra, MR; Schmelzle, T; Ladevèze, S; Kay, RF
    © 2014, Akademie der Naturwissenschaften Schweiz (SCNAT). During the Cenozoic paucituberculatans were much more diverse taxonomically and ecomorphologically than the three extant genera of shrew-like marsupials. Among paucituberculatans, palaeothentids were abundant during the Early Miocene, although most of the fossil remains consist of isolated teeth or fragmentary jaws. We describe a new and exceptional partial skeleton of Palaeothentes lemoinei (Palaeothentidae), collected from the Santa Cruz Formation (Santacrucian age, Early Miocene) in Patagonia. Whereas the skull of P. lemoinei has more plesiomorphic traits in the face, palate, and cranial vault than that of living paucituberculatans, the dental morphology is more derived. The osseous inner ear was examined using micro-CT scanning, revealing a cochlea with 1.9 turns, the presence of a “second crus commune”, an anterior semicircular canal (SC) projecting slightly dorsally from the dorsal-most point of the posterior SC, and lateral and posterior SCs projecting laterally to the same level. On the basis of postcranial anatomy, previous studies have demonstrated that P. lemoinei was an agile cursorial form, an inference supported by study of the new postcranial elements.
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    An improved approach to age-modeling in deep time: Implications for the Santa Cruz Formation, Argentina
    (Bulletin of the Geological Society of America, 2020-01-01) Trayler, RB; Schmitz, MD; Cuitiño, JI; Kohn, MJ; Bargo, MS; Kay, RF; Strömberg, CAE; Vizcaíno, SF
    © 2019 Geological Society of America. Accurate age-depth models for proxy records are crucial for inferring changes to the environment through space and time, yet traditional methods of constructing these models assume unrealistically small age uncertainties and do not account for many geologic complexities. Here we modify an existing Bayesian age-depth model to foster its application for deep time U-Pb and 40Ar/39Ar geochronology. More flexible input likelihood functions and use of an adaptive proposal algorithm in the Markov Chain Monte Carlo engine better account for the age variability often observed in magmatic crystal populations, whose dispersion can reflect inheritance, crystal residence times and daughter isotope loss. We illustrate this approach by calculating an age-depth model with a contiguous and realistic uncertainty envelope for the Miocene Santa Cruz Formation (early Miocene; Burdigalian), Argentina. The model is calibrated using new, high-precision isotope dilution U-Pb zircon ages for stratigraphically located interbedded tuffs, whose weighted mean ages range from ca. 16.78 ± 0.03 Ma to 17.62 ± 0.03 Ma. We document how the Bayesian age-depth model objectively reallocates probability across the posterior ages of dated horizons, and thus produces better estimates of relative ages among strata and variations in sedimentation rate. We also present a simple method to propagate age-depth model uncertainties onto stratigraphic proxy data using a Monte Carlo technique. This approach allows us to estimate robust uncertainties on isotope composition through time, important for comparisons of terrestrial systems to other proxy records.
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    Analysis of the Early-Middle Miocene mammal associations at the Río Santa Cruz (Patagonia, Argentina)
    (Publicacion Electronica de la Asociacion Paleontologica Argentina, 2019-01-01) Fernicola, JC; Vizcaíno, SF; Susana Bargo, M; Kay, RF; Cuitiño, JI
    © 2019 Asociacion Paleontologica Argentina. All rights reserved. The Santa Cruz Formation (SCF) records high latitude terrestrial paleoecosystems in the Southern Hemisphere during Burdigalian-early Langhian times (Early-Middle Miocene). Mammalian fossils from Río Santa Cruz (RSC) localities were first collected in the late 19th century, forming the basis for the Santacrucian South American Land Mammal Age. New collections permitt an update of the SCF mammalian species along the RSC. The total taxonomic richness is 95 mammalian species. Many species considered by Ameghino as exclusive for the older Notohippidian stage at similar latitude in the west, are not in fact so. The taxonomic richness in three localities along the RSC is substantially different: 47 species from Barrancas Blancas (BB), 60 from Segundas Barrancas Blancas (SBB) and nine from Yaten Huageno. The faunal composition between BB and SBB is also different: they share 31 species, of which six are present only at BB and 20 only at SBB. More than 85 % of all RSC species are also found at Atlantic coastal exposures of the SCF. In spite of BB (~17.04-16.49 Ma) being closer in age to coastal exposures, and SBB fossils (~16.46-15.63 Ma) being younger than the coastal localities (~17.80-16.30 Ma), the greatest similarity is between SBB and the coast. Faunal differences among the localities may be accounted for local variation in climatic and environmental factors. Previously proposed Santacrucian biozones should be set aside. The exposures of the SCF along the RSC should be considered as the type area of this unit and the Santacrucian fauna.
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    Auditory morphology and hearing sensitivity in fossil New World monkeys.
    (Anatomical record (Hoboken, N.J. : 2007), 2010-10) Coleman, M; Kay, RF; Colbert, MW
    In recent years it has become possible to investigate the hearing capabilities in fossils by analogy with studies in living taxa that correlate the bony morphology of the auditory system with hearing sensitivity. In this analysis, we used a jack-knife procedure to test the accuracy of one such study that examined the functional morphology of the primate auditory system and we found that low-frequency hearing (sound pressure level at 250 Hz) can be predicted with relatively high confidence (±3-8 dB depending on the structure). Based on these functional relationships, we then used high-resolution computed tomography to examine the auditory region of three fossil New World monkeys (Homunculus, Dolicocebus, and Tremacebus) and compared their morphology and predicted low-frequency sensitivity with a phylogenetically diverse sample of extant primates. These comparisons reveal that these extinct taxa shared many auditory characteristics with living platyrrhines. However, the fossil with the best preserved auditory region (Homunculus) also displayed a few unique features such as the relative size of the tympanic membrane and stapedial footplate and the degree of trabeculation of the anterior accessory cavity. Still, the majority of evidence suggests that these fossil species likely had similar low-frequency sensitivity to extant South American monkeys. This research adds to the small but growing body of evidence on the evolution of hearing abilities in extinct taxa and lays the groundwork for predicting hearing sensitivity in additional fossil primate specimens.
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    Fossil localities of the Santa Cruz Formation (Early Miocene, Patagonia, Argentina) prospected by Carlos Ameghino in 1887 revisited and the location of the Notohippidian
    (Journal of South American Earth Sciences, 2014-01-01) Fernicola, JC; Cuitiño, JI; Vizcaíno, SF; Bargo, MS; Kay, RF
    Between January and September of 1887 Carlos Ameghino carried out his first geologic and paleontological expedition to the Río Santa Cruz, Patagonia. Based on the fossils and geologic information compiled, in 1887 and 1889, Florentino Ameghino named more than 120 new species of extinct mammals and his Formación Santacruceña and Piso Santacruceño (Santacrucian stage). Data published by both brothers state that the specimens were collected in outcrops by the Río Santa Cruz, between 90 and 200km west of its mouth. However, information in the posthumously published letters and Travel Diary of C. Ameghino allows us to recognize a fourth locality, Río Bote, at about 50km further southwest. In 1900, 1902, F. Ameghino divided the Piso Santacruceño in a younger étage Santacruzienne and older étage Notohippidéen, restricting the geographical distribution of the latter to Kar Aiken locality, northeast of Lago Argentino. However, 15 of the 54 species that F. Ameghino listed as exclusively Notohippidian stage already had been named on specimens collected South to the Río Santa Cruz in 1887, two year prior to C. Ameghino's first visit to Kar Aiken. Based on historical information and several expeditions to the Río Santa Cruz and its environs, in this contribution we establish the geographical locations of the 1887 localities, formalize their names, evaluate the stratigraphic position of the fossil-bearing levels, and analyze the geographic extension of the Notohippidian, inferring that Río Bote is where C. Ameghino first collected species that came to define the Notohippidian. © 2014 Elsevier Ltd.
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    Fossil Localities Of The Santa Cruz Formation (Lower Miocene, Patagonia, Argentina) Prospected By Carlos Ameghino In 1887. The Problem Of The Notohippidian Stage
    (South American Journal of Earth Sciences, 2014) Fernicola, JC; Cuitiño, JI; Vizcaíno, SF; Bargo, MS; Kay, RF
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    Fossil vertebrates of the early-middle Miocene Cerro Boleadoras Formation, northwestern Santa Cruz Province, Patagonia, Argentina
    (Andean Geology, 2022-09-01) Vizcaíno, SF; Bargo, MS; Pérez, ME; Aramendía, I; Cuitiño, JI; Monsalvo, ES; Vlachos, E; Noriega, JI; Kay, RF
    The early-middle Miocene continental Cerro Boleadoras Formation (CBF) crops out in the area of Cerro Boleadoras and Cerro Plomo on the western slope of the Meseta del Lago Buenos Aires, northwestern Santa Cruz Province, Argentina. The lower levels of the CBF consist of laterally extensive medium to pebbly sandstone beds with trough cross-bedding, interpreted as fluvial channel deposits, interbedded with tabular fine-grained floodplain deposits. Recent fieldwork provided fossil vertebrates from these levels with an estimated age between ~16.5 Ma and 15.1 Ma (late Burdigalian-early Langhian). The studied section temporally overlaps with the middle or upper sections of the Santa Cruz Formation (SCF) in the Austral-Magallanes Basin of southern Patagonia, the Río Frías Formation in Chile, and the lower Collón Curá Formation of northern Patagonia. We compile an integrated faunal list for this locality, including specimens from previous collections, and discuss its chronological and paleoenvironmental implications. The taxa list includes most of the groups recorded in the SCF: one anuran, three birds, and at least 33 mammals (metatherians, xenarthrans, litopterns, notoungulate typotheres and caviomorph rodents), indicating a Santacrucian age sensu lato. We also recorded a testudine, which constitutes the southernmost record of tortoises in South America and worldwide. Faunal dissimilarities between the vertebrate fossil content of the CBF and the mentioned sections of the Santa Cruz, Río Frías and Collón Curá formations may reflect ecologic, climatic and geographic differences rather than temporal ones. The co-occurrence of arboreal or semiarboreal, browsing, frugivorous, and grazing mammals suggests the presence of both forested and open environments for the area occupied by the CBF rocks. However, it is not possible to discern whether these two environments coexisted or alternated, and whether one environment predominated over the other. Marker taxa, such as the chinchillid rodents Prolagostomus and Pliolagostomus, and the typothere Pachyrukhos indicate a trend to aridification during the Miocene in southern Patagonia, as previously reported for the upper part of the SCF along the Río Santa Cruz and south to the Río Coyle, along the Atlantic coast and the Río Gallegos.
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    HISTORICAL BACKGROUND FOR A REVISION OF THE PALEONTOLOGY OF THE SANTA CRUZ FORMATION (EARLY-MIDDLE MIOCENE) ALONG THE RÍO SANTA CRUZ, PATAGONIA, ARGENTINA
    (Publicación Electrónica de la Asociación Paleontológica Argentina, 2020) Fernicola, JC; Bargo, MS; Vizcaíno, SF; Kay, RF
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    Insights on the controls on floodplain-dominated fluvial successions: A perspective from the early–middle miocene santa cruz formation in río chalía (patagonia, argentina)
    (Journal of the Geological Society, 2021-01-01) Cuitiño, JI; Raigemborn, MS; Bargo, MS; Vizcaíno, SF; Muñoz, NA; Kohn, MJ; Kay, RF
    The Santa Cruz Formation (SCF) in Río Chalía (Austral Basin, Patagonia, Argentina) is a well-exposed fluvial succession with abundant and diverse fossil vertebrates accumulated during the Miocene Climatic Optimum (MCO). Using facies analysis, characterization of stratigraphic architecture, U–Pb geochronology and vertebrate palaeontology, we assess the timing and interplay of controlling factors on the sedimentation, including tectonics, global sea level, climate and sediment supply. Throughout the succession, there occurred a constant aggradation of the floodplain-dominated fluvial system. Seven zircon U–Pb ages constrain the time of accumulation between c. 18 and 15.2 Ma, under a relatively constant sedimentation rate of 150 ± 50 m myr–1 . The large number of fossil vertebrates indicates a Santacrucian fauna, showing no recognizable changes through the section. The basin-scale, low-gradient anastomosed fluvial system of the SCF records a period of about 3 myr of relatively constant environmental conditions controlled by continuous basin subsidence and high sediment supply conditioned by explosive volcanism together with weathering of uplifting terrains in the Andes. In addition, the system was influenced by a temperate to warm and subhumid climate favoured by the MCO before the onset of the Andean rain shadow, together to high global sea levels.
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    Introducing molaR: a New R Package for Quantitative Topographic Analysis of Teeth (and Other Topographic Surfaces)
    (Journal of Mammalian Evolution, 2016-12-01) Pampush, JD; Winchester, JM; Morse, PE; Vining, AQ; Boyer, DM; Kay, RF
    © 2016, Springer Science+Business Media New York.Researchers studying mammalian dentitions from functional and adaptive perspectives increasingly have moved towards using dental topography measures that can be estimated from 3D surface scans, which do not require identification of specific homologous landmarks. Here we present molaR, a new R package designed to assist researchers in calculating four commonly used topographic measures: Dirichlet Normal Energy (DNE), Relief Index (RFI), Orientation Patch Count (OPC), and Orientation Patch Count Rotated (OPCR) from surface scans of teeth, enabling a unified application of these informative new metrics. In addition to providing topographic measuring tools, molaR has complimentary plotting functions enabling highly customizable visualization of results. This article gives a detailed description of the DNE measure, walks researchers through installing, operating, and troubleshooting molaR and its functions, and gives an example of a simple comparison that measured teeth of the primates Alouatta and Pithecia in molaR and other available software packages. molaR is a free and open source software extension, which can be found at the doi:10.13140/RG.2.1.3563.4961 (molaR v. 2.0) as well as on the Internet repository CRAN, which stores R packages.
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    Leonard B. Radinsky (1937–1985), Radical Biologist
    (Journal of Mammalian Evolution, 2019-01-01) Kay, RF
    © 2019, Springer Science+Business Media, LLC, part of Springer Nature. Trained in vertebrate paleontology, Leonard Radinsky (1937–1985) made signal contributions to the study of form and function in paleobiology. Here, I review Radinsky’s contributions and philosophy in the context of developments during the 1960s and 1970s, when a significant number of vertebrate paleontologists departed from their roots in the geological sciences to embrace a new interest in paleobiology and evolution. The study of comparative biomechanics and allometry in extant mammals was brought to the fore, with the express intent of applying the findings to reconstruct the biology of their extinct relatives. Radinsky’s contributions lay especially in the area of jaw mechanics in carnivorans and ungulates, and the evolution of the brain in ungulates, carnivorans, and primates. Alongside his important scientific contributions, Radinsky espoused radical views for his time. He fervently believed that basic science cannot be isolated from its social and political context. At a time when the US was deeply engaged in military conflict in Southeast Asia, Radinsky believed that the results of basic science unjustly were being co-opted by corporate and military interests. He believed that science should be used for the betterment of the great majority of the people.
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    Mammalian faunas, ecological indices, and machine-learning regression for the purpose of paleoenvironment reconstruction in the Miocene of South America
    (Palaeogeography, Palaeoclimatology, Palaeoecology, 2019-03-15) Spradley, JP; Glazer, BJ; Kay, RF
    © 2019 Elsevier B.V. Reconstructing paleoenvironments has long been considered a vital component for understanding community structure of extinct organisms, as well as patterns that guide evolutionary pathways of species and higher-level taxa. Given the relative geographic and phylogenetic isolation of the South American continent throughout much of the Cenozoic, the South American fossil record presents a unique perspective of mammalian community evolution in the context of changing climates and environments. Here we focus on one line of evidence for paleoenvironment reconstruction: ecological diversity, i.e. the number and types of ecological niches filled within a given fauna. We propose a novel approach by utilizing ecological indices as predictors in two regressive modeling techniques—Random Forest (RF) and Gaussian Process Regression (GPR)—which are applied to 85 extant Central and South American localities to produce paleoecological prediction models. Faunal richness is quantified via ratios of ecologies within the mammalian communities, i.e. ecological indices, which serve as predictor variables in our models. Six climate/habitat variables were then predicted using these ecological indices: mean annual temperature (MAT), mean annual precipitation (MAP), temperature seasonality, precipitation seasonality, canopy height, and net primary productivity (NPP). Predictive accuracy of RF and GPR is markedly higher when compared to previously published methods. MAT, MAP, and temperature seasonality have the lowest predictive error. We use these models to reconstruct paleoclimatic variables in two well-sampled Miocene faunas from South America: fossiliferous layers (FL) 1–7, Santa Cruz Formation (Early Miocene), Santa Cruz Province, Argentina; and the Monkey Beds unit, Villavieja Formation (Middle Miocene) Huila, Colombia. Results suggest general concordance with published estimations of precipitation and temperature, and add information with regards to the other climate/habitat variables included here. Ultimately, we believe that RF and GPR in conjunction with ecological indices have the potential to contribute to paleoenvironment reconstruction.
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    New primates from the Río Santa Cruz and Río Bote (Early-Middle Miocene), Santa Cruz Province, Argentina
    (Publicacion Electronica de la Asociacion Paleontologica Argentina, 2019-01-01) Kay, RF; Perry, JMG
    © 2019 Asociacion Paleontologica Argentina. All rights reserved. Four specimens of primates were collected from the Santa Cruz Formation (Early-Middle Miocene) during expeditions undertaken by the Museo de la Plata, Museo Argentino de Ciencias Naturales “Bernardino Rivadavia”, and Duke University in 2013, 2014, and 2017. A new species of Homunculus Ameghino, H. vizcainoi (Platyrrhini, Homunculidae), was identified at Barrancas Blancas, and Segundas Barrancas Blancas localities on the right bank of the Río Santa Cruz (Santa Cruz Province, Argentina). The Barrancas Blancas specimen comes from a tuff dated at 17.04 Ma; those from Segundas Barrancas Blancas are older than a tuff dated at 16.32 Ma and younger than a tuff dated at 17.36 Ma. A Río Bote specimen is confidently identified as Homunculus, but of uncertain species. All these fossil primates are temporally equivalent to those from the coastal Santa Cruz Formation, and younger than those from the Pinturas Formation to the north. By contrast, the lower and middle strata of the Pinturas Formation contain a different but closely related taxon, Carlocebus Fleagle. All known records of Carlocebus from the Pinturas Formation in north central Santa Cruz Province are older than the known occurrences of Homunculus in the Santa Cruz Formation in the Río Santa Cruz valley, Río Bote and elsewhere.
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    Paleoenvironmental reconstruction of the coastal Monte Léon and Santa Cruz formations (Early Miocene) at Rincón del Buque, Southern Patagonia: A revisited locality
    (Journal of South American Earth Sciences, 2015-07-01) Raigemborn, MS; Matheos, SD; Krapovickas, V; Vizcaíno, SF; Bargo, MS; Kay, RF; Fernicola, JC; Zapata, L
    © 2015 Elsevier Ltd.Sedimentological, ichnological and paleontological analyses of the Early Miocene uppermost Monte León Formation and the lower part of the Santa Cruz Formation were carried out in Rincón del Buque (RDB), a fossiliferous locality north of Río Coyle in Santa Cruz Province, Patagonia, Argentina. This locality is of special importance because it contains the basal contact between the Monte Léon (MLF) and the Santa Cruz (SCF) formations and because it preserves a rich fossil assemblage of marine invertebrates and marine trace fossils, and terrestrial vertebrates and plants, which has not been extensively studied. A ~90m-thick section of the MLF and the SCF that crops out at RDB was selected for this study. Eleven facies associations (FA) are described, which are, from base to top: subtidal-intertidal deposits with Crassotrea orbignyi and bioturbation of the Skolithos-Cruziana ichnofacies (FA1); tidal creek deposits with terrestrial fossil mammals and Ophiomorpha isp. burrows (FA2); tidal flat deposits with Glossifungites ichnofacies (FA3); deposits of tidal channels (FA4) and tidal sand flats (FA5) both with and impoverish Skolithos ichnofacies associated; marsh deposits (FA6); tidal point bar deposits recording a depauperate mixture of both the Skolithos and Cruziana ichnofacies (FA7); fluvial channel deposits (FA8); fluvial point bar deposits (FA9); floodplain deposits (FA10); and pyroclastic and volcaniclastic deposits of the floodplain where terrestrial fossil mammal remains occur (FA11).The transition of the MLF-SCF at RDB reflects a changing depositional environment from the outer part of an estuary (FA1) through the central (FA2-6) to inner part of a tide-dominated estuary (FA7). Finally a fluvial system occurs with single channels of relatively low energy and low sinuosity enclosed by a broad, low-energy floodplain dominated by partially edaphized ash-fall, sheet-flood, and overbank deposits (FA8-11). Pyroclastic and volcaniclastic materials throughout the succession must have been deposited as ash-fall distal facies in a fluvial setting and also were carried by fluvial streams and redeposited in both estuarine and fluvial settings. These materials preserve most of the analyzed terrestrial fossil mammals that characterize the Santacrucian age of the RDB's succession. Episodic sedimentation under volcanic influence, high sedimentation rates and a relatively warm and seasonal climate are inferred for the MLF and SCF section.Lateral continuity of the marker horizons at RDB serve for correlation with other coastal localities such as the lower part of the coastal SCF south of Río Coyle (~17.6-17.4Ma) belonging to the Estancia La Costa Member of the SCF.
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    Paleoenvironments and paleoecology of the Santa Cruz Formation (early-middle Miocene) along the Río Santa Cruz, Patagonia (Argentina)
    (Journal of South American Earth Sciences, 2021-08) Kay, RF; Vizcaíno, SF; Bargo, MS; Spradley, JP; Cuitiño, JI